Solanaceae

<h2 id="etymology">Etymology</h2>
The name "Solanaceae" comes from <a href="/facts/Solanum/UDkyXGxg">Solanum</a>, the <a href="/facts/Type_genus/vKbAuKph">type genus</a> of the family, + -aceae,<a class="footnote-ref" id="fnref:2" href="#fn:2">2</a> a standardized <a href="/facts/Suffix/Nlnft3NK">suffix</a> for plant family names in modern taxonomy. The etymology of the word solanum is unclear. The name probably comes from a perceived resemblance of certain species' flowers to the sun (sol in <a href="/facts/Latin/pM3Kge85">Latin</a>) and its rays. At least one species of Solanum is known as the "sunberry". Alternatively, the name could originate from the Latin verb solare, meaning "to soothe", presumably referring to the soothing <a href="/facts/Pharmacology/4Rx8s7Ie">pharmacological</a> properties of some of the <a href="/facts/Psychoactive/dNXrUmsx">psychoactive</a> species of the family.<a class="footnote-ref" id="fnref:3" href="#fn:3">3</a>
The common name "nightshade" developed directly from <a href="/facts/Middle_English/dj1wHvwd">Middle English</a> nyght-shade, originating from the <a href="/facts/Old_English/9rsmQG2a">Old English</a> word nihtscada (lit. "shade of night"). <a href="/facts/Cognate/nXETf5Y4">Cognates</a> are found in many <a href="/facts/Germanic_languages/BxAJX3Kf">Germanic languages</a>, such as the <a href="/facts/German_language/oWaQIDA8">German</a> word nachtschatten and the <a href="/facts/Dutch_language/yIB5lbsx">Dutch</a> word nachtschade. The reason for the usage of the name is ultimately unknown, but it could have been a reference to the appearance of the fruits.<a class="footnote-ref" id="fnref:4" href="#fn:4">4</a>

<h2 id="description">Description</h2>

Nightshades can take the form of herbs, <a href="/facts/Shrub/X1gKVxL5">shrubs</a>, <a href="/facts/Tree/3q0G4cbP">trees</a>, <a href="/facts/Vine/3nVRIAws">vines</a> and lianas, and sometimes <a href="/facts/Epiphytes/n0oUN0Yd">epiphytes</a>. They can be <a href="/facts/Annual_plant/SkPv72DA">annuals</a>, <a href="/facts/Biennial_plant/utHmQsd6">biennials</a>, or <a href="/facts/Perennial_plant/MQ3NvAUW">perennials</a>, upright or decumbent. Some have subterranean <a href="/facts/Tuber/wGw2Sbag">tubers</a>. They do not have <a href="/facts/Laticifer/w62kHnjS">laticifers</a>, nor <a href="/facts/Latex/c1lvjF0A">latex</a>, nor coloured <a href="/facts/Sap/cxdgQYP8">saps</a>.
They can have a basal or terminal group of leaves or neither of these types. The <a href="/facts/Leaf/1m6wCNTg">leaves</a> are generally alternate or alternate to opposed (that is, alternate at the base of the plant and opposed towards the <a href="/facts/Inflorescence/CvpNd4vS">inflorescence</a>). The leaves can be herbaceous, leathery, or transformed into <a href="/facts/Thorns%2c_spines_and_prickles/gS6HPTqb">spines</a>. The leaves are generally <a href="/facts/Petiole_(botany)/pext65Wr">petiolate</a> or subsessile, rarely sessile. They are frequently inodorous, but some are aromatic or fetid. The foliar lamina can be either simple or compound, and the latter can be either pinnatifid or ternate. The leaves have reticulated venation and lack a basal <a href="/facts/Meristem/YlMKp2W5">meristem</a>. The laminae are generally dorsiventral and lack secretory cavities. The <a href="/facts/Stoma/Rer9Nzct">stomata</a> are generally confined to one of a leaf's two sides; they are rarely found on both sides.

The <a href="/facts/Flower/7gI5Kefw">flowers</a> are generally <a href="/facts/Hermaphrodite/C4aYIk4n">hermaphrodites</a>, although some are <a href="/facts/Monoecy/Up20kQT3">monoecious</a>, <a href="/facts/Andromonoecy/sy4U3zS6">andromonoecious</a>, or <a href="/facts/Dioecy/jwMHLkuM">dioecious</a> species (such as some Solanum or Symonanthus). They are most commonly <a href="/facts/Pollination/u383g7XJ">pollinated</a> by <a href="/facts/Entomophily/cSwptQsL">insects</a>.<a class="footnote-ref" id="fnref:5" href="#fn:5">5</a> The flowers can be solitary or grouped into terminal, cymose, or axillary inflorescences. The flowers are medium-sized, fragrant (Nicotiana), fetid (Anthocercis), or inodorous. The flowers are usually <a href="/facts/Floral_symmetry/oJMuxKtL">actinomorphic</a>, slightly <a href="/facts/Zygomorphic/oJMuxKtL">zygomorphic</a>, or markedly zygomorphic (for example, in flowers with a bilabial corolla in Schizanthus species). The irregularities in symmetry can be due to the <a href="/facts/Androecium/MzNPVK3D">androecium</a>, to the <a href="/facts/Perianth/dy8xGxeb">perianth</a>, or both at the same time. In the great majority of species, the flowers have a differentiated perianth with a calyx and <a href="/facts/Corolla_(flower)/DQMkpRSm">corolla</a> (with five sepals and five petals, respectively) an androecium with five <a href="/facts/Stamen/MzNPVK3D">stamens</a> and two <a href="/facts/Carpel/P5qFQMk1">carpels</a> forming a <a href="/facts/Gynoecium/P5qFQMk1">gynoecium</a> with a superior <a href="/facts/Ovary_(botany)/Qug4yDY4">ovary</a><a class="footnote-ref" id="fnref:6" href="#fn:6">6</a> (they are therefore referred to as pentamers and tetracyclic). The <a href="/facts/Stamen/MzNPVK3D">stamens</a> are <a href="/facts/Stamen/MzNPVK3D">epipetalous</a> and are typically present in multiples of four or five, most commonly four or eight. They usually have a hypogynous disk. The calyx is gamosepalous (as the sepals are joined forming a tube), with the (4)5(6) segments equal, it has five lobes, with the lobes shorter than the tube, it is persistent and often accrescent. The corolla usually has five petals that are also joined forming a tube. Flower shapes are typically rotate (wheel-shaped, spreading in one plane, with a short tube) or tubular (elongated cylindrical tube), campanulated, or funnel-shaped.
The androecium has (2)(4)5(6) free stamens within its opposite sepals (they alternate with the petals). They are usually fertile or, in some cases (for example in Salpiglossideae) they have <a href="/facts/Staminode/44XDV0IQ">staminodes</a>. In the latter case, there is usually either one staminode (Salpiglossis) or three (Schizanthus). The anthers touch on their upper end forming a ring, or they are completely free, dorsifixed, or basifixed with poricide dehiscence or through small longitudinal cracks. The stamen's <a href="/facts/Filament_(botany)/MzNPVK3D">filament</a> can be filiform or flat. The stamens can be inserted inside the coralline tube or exserted. The plants demonstrate simultaneous microsporogenesis, the microspores are tetrad, tetrahedral, or isobilateral. The pollen grains are bicellular at the moment of dehiscence, usually open and angular.
The gynoecium is bicarpelar (rarely three- or five-locular) with a <a href="/facts/Hypogynous/Qug4yDY4">superior</a> <a href="/facts/Ovary_(plants)/Qug4yDY4">ovary</a> and two <a href="/facts/Locule/0j8YPj1q">locules</a>, which may be secondarily divided by false <a href="/facts/Septum/lPNxGG9u">septa</a>, as is the case for Nicandreae and Datureae. The gynoecium is located in an oblique position relative to the flower's median plane. They have one <a href="/facts/Style_(botany)/JEQ85ChP">style</a> and one <a href="/facts/Stigma_(botany)/JEQ85ChP">stigma</a>; the latter is simple or bilobate. Each locule has one to 50 ovules that are anatropous or hemianatropous with axillar placentation. The development of the <a href="/facts/Ovule/DFvS1VvA">embryo sack</a> can be the same as for Polygonum or Allium species. The embryo sack's nuclear poles become fused before <a href="/facts/Fertilization/g8u3UKzF">fertilization</a>. The three antipodes are usually ephemeral or persistent as in the case of Atropa. The <a href="/facts/Fruit/PBpuzk60">fruit</a> can be a <a href="/facts/Berry_(botany)/2JfhI0vw">berry</a> as in the case of the tomato or wolfberry, or a <a href="/facts/Dehiscence_(botany)/w5cbltMG">dehiscent</a> <a href="/facts/Capsule_(fruit)/nSgcw6KK">capsule</a> as in Datura, or a <a href="/facts/Drupe/WrnGMPSa">drupe</a>. The fruit has <a href="/facts/Placentation/nqcxcaPM">axial placentation</a>. The capsules are normally septicidal or rarely loculicidal or valvate. The <a href="/facts/Seed/a4xGSLxJ">seeds</a> are usually endospermic, oily (rarely starchy), and without obvious hairs. The seeds of most Solanaceae are round and flat, about 2–4 mm (0.079–0.157 in) in diameter. The embryo can be straight or curved, and has two cotyledons. Most species in the Solanaceae have 2n=24 <a href="/facts/Chromosome/FRGf11FC">chromosomes</a>,<a class="footnote-ref" id="fnref:7" href="#fn:7">7</a> but the number may be a higher multiple of 12 due to <a href="/facts/Polyploidy/L7xyKHdK">polyploidy</a>. Wild <a href="/facts/Potato/F8jRVExs">potatoes</a>, of which there are about 200, are predominantly diploid (2 × 12 = 24 chromosomes), but triploid (3 × 12 = 36 chromosomes), tetraploid (4 × 12 = 48 chromosomes), pentaploid (5 × 12 = 60) and even hexaploid (6 × 12 = 72 chromosome) species or populations exist. The cultivated species Solanum tuberosum has 4 × 12 = 48 chromosomes. Some Capsicum species have 2 × 12 = 24 chromosomes, while others have 26 chromosomes.

<h3>Diversity of characteristics</h3>
Despite the previous description, the Solanaceae exhibit a large morphological variability, even in their reproductive characteristics. Examples of this diversity include:<a class="footnote-ref" id="fnref:8" href="#fn:8">8</a><a class="footnote-ref" id="fnref:9" href="#fn:9">9</a>

<ul><li>The number of carpels that form the gynoecium</li></ul>
In general, the Solanaceae have a gynoecium (the female part of the flower) formed of two carpels. However, Melananthus has a monocarpelar gynoecium, there are three or four carpels in <a href="/facts/Capsicum/oqoEFu4F">Capsicum</a>, three to five in <a href="/facts/Nicandra/sA4LsruE">Nicandra</a>, some species of <a href="/facts/Jaborosa/Y0qHgouq">Jaborosa</a> and <a href="/facts/Trianaea/StSTIj16">Trianaea</a> and four carpels in Iochroma umbellatum.

<ul><li>The number of locules in the ovary</li></ul>
The number of locules in the ovary is usually the same as the number of carpels. However, some species occur in which the numbers are not the same due to the existence of false septa (internal walls that subdivide each locule), such as in Datura and some members of the Lycieae (the genera <a href="/facts/Grabowskia/wqlIxG6N">Grabowskia</a> and Vassobia).

<ul><li>Type of ovules and their number</li></ul>
The ovules are generally inverted, folded sharply backwards (anatropous), but some genera have ovules that are rotated at right angles to their stalk (campilotropous) as in Phrodus, <a href="/facts/Grabowskia/wqlIxG6N">Grabowskia</a> or Vassobia), or are partially inverted (hemitropous as in <a href="/facts/Cestrum/YbBzKxS5">Cestrum</a>, Capsicum, <a href="/facts/Schizanthus/n3RYlUYF">Schizanthus</a> and Lycium). The number of ovules per locule also varies from a few (two pairs in each locule in Grabowskia, one pair in each locule in Lycium) and very occasionally only one ovule is in each locule as for example in Melananthus.

<ul><li>The type of fruit</li></ul>
The fruits of the great majority of the Solanaceae are berries or capsules (including pyxidia) and less often drupes.
Berries are common in the subfamilies Cestroideae, Solanoideae (with the exception of Datura, Oryctus, Grabowskia and the tribe Hyoscyameae) and the tribe Juanulloideae (with the exception of <a href="/facts/Markea/njKCohtd">Markea</a>).
Capsules are characteristic of the subfamilies Cestroideae (with the exception of <a href="/facts/Cestrum/YbBzKxS5">Cestrum</a>) and Schizanthoideae, the tribes Salpiglossoideae and Anthocercidoideae, and the genus Datura. The tribe Hyoscyameae has pyxidia.
Drupes are typical of the Lycieae tribe and in Iochrominae.<a class="footnote-ref" id="fnref:10" href="#fn:10">10</a>

<h2 id="taxonomy">Taxonomy</h2>

The first scientific reference to Solanaceae was in 1763 in French naturalist <a href="/facts/Michel_Adanson/G7eWPH8l">Michel Adanson</a>'s Familles des Plantes.<a class="footnote-ref" id="fnref:11" href="#fn:11">11</a> He did not use a formal name for his taxon, and simply labeled it as "Les Solanum".<a class="footnote-ref" id="fnref:12" href="#fn:12">12</a>: 215  He included a total of 10 genera, all of which are still accepted as members of the family.<a class="footnote-ref" id="fnref:13" href="#fn:13">13</a>: 218–219 <a class="footnote-ref" id="fnref:14" href="#fn:14">14</a> Adanson is however not considered to be the authority of the family, that title instead being held by French botanist <a href="/facts/Antoine_Laurent_de_Jussieu/Ip3CSVvv">Antoine Laurent de Jussieu</a>, who gave the group a formal scientific name in 1789 in his Genera Plantarum [fr].<a class="footnote-ref" id="fnref:15" href="#fn:15">15</a> Jussieu classified the taxon as an <a href="/facts/Order_(biology)/GKLJvIpt">order</a> and used the name "Solaneæ". His order included 19 genera, some of which—such as <a href="/facts/Verbascum_blattaria/AcI6gFGy">Verbascum blattaria</a>,<a class="footnote-ref" id="fnref:16" href="#fn:16">16</a> <a href="/facts/Avicennia/g06lGnzF">Bontia</a>,<a class="footnote-ref" id="fnref:17" href="#fn:17">17</a> and <a href="/facts/Crescentia_cujete/oUqIlPSX">Crescentia cujete</a><a class="footnote-ref" id="fnref:18" href="#fn:18">18</a>—are no longer considered members of the family.<a class="footnote-ref" id="fnref:19" href="#fn:19">19</a>: 124–127  Some genera Jussieu included within Solanaceae he proposed could belong to <a href="/facts/Boraginaceae/wGbPMTXF">Boraginaceae</a>, which he also named in Genera Plantarum; he noted that there was a high degree of similarity between Solanaceae and Boraginaceae, and proposed that they could be considered one order.<a class="footnote-ref" id="fnref:20" href="#fn:20">20</a>: 132 
Following Jussieu's publication, taxonomists have heavily revised, re-examined, and added to the taxon. "Solaneæ" was reclassified as a <a href="/facts/Family_(biology)/qTrn6E8E">family</a> by the 1820s,<a class="footnote-ref" id="fnref:21" href="#fn:21">21</a>: 325  and began to be called "Solanaceae" by some authors around the 1830s,<a class="footnote-ref" id="fnref:22" href="#fn:22">22</a> which became the standard name by 1905 per the <a href="/facts/International_Code_of_Nomenclature_for_algae%2c_fungi%2c_and_plants/EbaUF9P5">ICBN</a> nomenclature rules.<a class="footnote-ref" id="fnref:23" href="#fn:23">23</a>
In 1835, <a href="/facts/Gilbert_Thomas_Burnett/TVdf7cNs">Gilbert Burnett</a> was the first to publish a subclassification of Solanaceae, and included 4 subgroups: <a href="/facts/Cestroideae/086ugKao">Cestridæ</a>, Nolanidæ, <a href="/facts/Solanoideae/4MwH5WzK">Solanidæ</a>, and Verbascidæ.<a class="footnote-ref" id="fnref:24" href="#fn:24">24</a>: 1106  While Cestridæ and Solanidæ were broadly accepted as subfamilies (later renamed Cestroideae and Solanoideae), Nolanidæ and Verbascidæ, having several non-solanaceous characteristics, were only tentatively assigned to the family by Burnett<a class="footnote-ref" id="fnref:25" href="#fn:25">25</a>: 987  and eventually were split from the family.<a class="footnote-ref" id="fnref:26" href="#fn:26">26</a> While several core genera were widely accepted to be a part of Solanaceae, others have been less stable in their placement. The problem of some species having a mix of solanaceous and non-solanaceous traits continued to be a significant source of conflict in Solanaceae taxnomy.<a class="footnote-ref" id="fnref:27" href="#fn:27">27</a><a class="footnote-ref" id="fnref:28" href="#fn:28">28</a> The families <a href="/facts/Duckeodendron/l9nItUbL">Duckeodendraceae</a>, Goetzaceae, and <a href="/facts/Nolana/93BmSNsj">Nolanaceae</a> were particularly tantalizing; it had long been known that they were closely related to Solanceae, but to what extent was unclear.<a class="footnote-ref" id="fnref:29" href="#fn:29">29</a><a class="footnote-ref" id="fnref:30" href="#fn:30">30</a> All three families have extremely similar wood anatomy to Solanaceae,<a class="footnote-ref" id="fnref:31" href="#fn:31">31</a> and in at least the case of Goetzeaceae leaf anatomy as well.<a class="footnote-ref" id="fnref:32" href="#fn:32">32</a>
The advent of <a href="/facts/Molecular_phylogenetics/FysplHP5">molecular phylogenetics</a> in the late 20th century allowed genetic and chemical data to be incorporated into <a href="/facts/Cladistics/JasKVABk">cladistics</a>, providing a new robust method of uncovering evolutionary relationships.<a class="footnote-ref" id="fnref:33" href="#fn:33">33</a> An early molecular study by Olmstead et al. in 1999 provided a significant update to Solanaceae taxonomy, splitting Cestroideae into 5 subfamilies, Solaneae into multiple tribes, and finding <a href="/facts/Nolana/93BmSNsj">Nolana</a> and Geotzeaceae (demoted to subfamily Geotzoideae) to indeed be members of the family.<a class="footnote-ref" id="fnref:34" href="#fn:34">34</a> Further studies found Duckeodendraceae to be in the family as well.<a class="footnote-ref" id="fnref:35" href="#fn:35">35</a> The contents of the family are now mostly agreed upon, although the exact position of the subgroups is still debated.<a class="footnote-ref" id="fnref:36" href="#fn:36">36</a> The December 2024 <a href="/facts/World_Flora_Online/fqY9CS1x">World Flora Online</a> classification lists 8 subfamilies, 18 tribes, 103 genera, and 2,729 species in the family, shown below.<a class="footnote-ref" id="fnref:37" href="#fn:37">37</a><a class="footnote-ref" id="fnref:38" href="#fn:38">38</a>

<table><tbody><tr><td>Subfamily Cestroideae<ul><li><a href="/facts/Protoschwenkia/gz4wwuDK">Protoschwenkia</a> (1 sp.)</li><li>Tribe Benthamiellae<ul><li><a href="/facts/Benthamiella/f5ptBtDy">Benthamiella</a> (12 sp.)</li><li><a href="/facts/Combera/6Mtur4Ru">Combera</a> (2 sp.)</li></ul></li><li>Tribe Browallieae<ul><li><a href="/facts/Browallia/YxF68PYA">Browallia</a> (22 sp.)</li><li><a href="/facts/Streptosolen/ox269EIz">Streptosolen</a> (1 sp.)</li></ul></li><li>Tribe Cestreae<ul><li><a href="/facts/Cestrum/YbBzKxS5">Cestrum</a> (172 sp.)</li><li><a href="/facts/Sessea/jPldBQjp">Sessea</a> (24 sp.)</li><li><a href="/facts/Vestia_foetida/QT9c2TFB">Vestia</a> (1 sp.)</li></ul></li><li>Tribe Salpiglossideae<ul><li><a href="/facts/Reyesia/vQYBmxYB">Reyesia</a> (4 sp.)</li><li><a href="/facts/Salpiglossis/PAtQ6uL6">Salpiglossis</a> (2 sp.)</li></ul></li></ul>Subfamily Duckeodendroideae<ul><li><a href="/facts/Duckeodendron/l9nItUbL">Duckeodendron</a> (1 sp.)</li></ul>Subfamily Goetzeoideae<ul><li><a href="/facts/Coeloneurum/ufx6JRQB">Coeloneurum</a> (1 sp.)</li><li><a href="/facts/Espadaea/kGxo1DPU">Espadaea</a> (1 sp.)</li><li><a href="/facts/Goetzea/tnFbpIoB">Goetzea</a> (2 sp.)</li><li><a href="/facts/Henoonia/GC2fGYvr">Henoonia</a> (1 sp.)</li><li><a href="/facts/Metternichia/vLBmkWzh">Metternichia</a> (1 sp.)</li><li><a href="/facts/Tsoala/KRaIJGf7">Tsoala</a> (1 sp.)</li></ul></td><td>Subfamily Nicotianoideae<ul><li>Tribe Anthocerideae<ul><li><a href="/facts/Anthocercis/HeDgx6d2">Anthocercis</a> (10 sp.)</li><li><a href="/facts/Anthotroche/1I2sXVJ4">Anthotroche</a> (3 sp.)</li><li><a href="/facts/Crenidium/SEAzp9y7">Crenidium</a> (1 sp.)</li><li><a href="/facts/Cyphanthera/D1jUBX3Q">Cyphanthera</a> (8 sp.)</li><li><a href="/facts/Duboisia/pcL4Xow7">Duboisia</a> (4 sp.)</li><li><a href="/facts/Grammosolen/LfzUUMq8">Grammosolen</a> (4 sp.)</li><li><a href="/facts/Symonanthus/lpeLpy88">Symonanthus</a> (2 sp.)</li></ul></li><li>Tribe Nicotianeae<ul><li><a href="/facts/Nicotiana/Bfh1JnE0">Nicotiana</a> (101 sp.)</li></ul></li></ul>Subfamily Petunioideae<ul><li>Tribe Petunieae<ul><li><a href="/facts/Bouchetia_(plant)/pHhhydbY">Bouchetia</a> (3 sp.)</li><li><a href="/facts/Brunfelsia/7M9CTD0b">Brunfelsia</a> (49 sp.)</li><li><a href="/facts/Calibrachoa/D4vvl22r">Calibrachoa</a> (29 sp.)</li><li><a href="/facts/Fabiana_(plant)/7Ty4F9Pg">Fabiana</a> (16 sp.)</li><li><a href="/facts/Hunzikeria/X0MzoQsS">Hunzikeria</a> (4 sp.)</li><li><a href="/facts/Leptoglossis/pWuo9Emr">Leptoglossis</a> (6 sp.)</li><li><a href="/facts/Nierembergia/73pDbu60">Nierembergia</a> (20 sp.)</li><li>Petchoa (hybrid)</li><li><a href="/facts/Petunia/ASe88e60">Petunia</a> (16 sp.)</li><li><a href="/facts/Plowmania/BvTuzEGr">Plowmania</a> (1 sp.)</li></ul></li></ul>Subfamily Schizanthoideae<ul><li><a href="/facts/Schizanthus/n3RYlUYF">Schizanthus</a> (16 sp.)</li></ul>Subfamily Schwenckioideae<ul><li><a href="/facts/Heteranthia/Rddg8uTz">Heteranthia</a> (1 sp.)</li><li><a href="/facts/Melananthus/ymdpdE1W">Melananthus</a> (5 sp.)</li><li><a href="/facts/Schwenckia/hNdXQCBI">Schwenckia</a> (20 sp.)</li></ul></td><td>Subfamily Solanoideae<ul><li><a href="/facts/Nectouxia/eL8Ikge6">Nectouxia</a> (1 sp.)</li><li><a href="/facts/Salpichroa/dlb2DgkQ">Salpichroa</a> (22 sp.)</li><li>Tribe Capsiceae<ul><li><a href="/facts/Capsicum/oqoEFu4F">Capsicum</a> (43 sp.)</li><li><a href="/facts/Lycianthes/7DBqj0XV">Lycianthes</a> (137 sp.)</li></ul></li><li>Tribe Datureae<ul><li><a href="/facts/Brugmansia/TD8YtCmd">Brugmansia</a> (10 sp.)</li><li><a href="/facts/Datura/ma7yqk7U">Datura</a> (15 sp.)</li><li><a href="/facts/Trompettia/EGfFdBSR">Trompettia</a> (1 sp.)</li></ul></li><li>Tribe Hyoscyameae<ul><li><a href="/facts/Anisodus/JmKMX2kh">Anisodus</a> (4 sp.)</li><li><a href="/facts/Atropa/1WlXXWgL">Atropa</a> (1 sp.)</li><li><a href="/facts/Atropanthe/hkp1m8CD">Atropanthe</a> (1 sp.)</li><li><a href="/facts/Hyoscyamus/JfYN6kbo">Hyoscyamus</a> (17 sp.)</li><li><a href="/facts/Physochlaina/MKP2SnGE">Physochlaina</a> (7 sp.)</li><li><a href="/facts/Przewalskia/Kjf0NUuQ">Przewalskia</a> (1 sp.)</li><li><a href="/facts/Scopolia/zavSULaE">Scopolia</a> (2 sp.)</li></ul></li><li>Tribe Jaboroseae<ul><li><a href="/facts/Jaborosa/Y0qHgouq">Jaborosa</a> (23 sp.)</li></ul></li><li>Tribe Latueae<ul><li><a href="/facts/Latua/IMpB0xXU">Latua</a> (1 sp.)</li></ul></li><li>Tribe Lycieae<ul><li><a href="/facts/Lycium/4VVD4sFu">Lycium</a> (93 sp.)</li><li><a href="/facts/Nolana/93BmSNsj">Nolana</a> (98 sp.)</li><li><a href="/facts/Sclerophylax/AQD3axm6">Sclerophylax</a> (12 sp.)</li></ul></li><li>Tribe Mandragoreae<ul><li><a href="/facts/Mandragora_(genus)/fVL6VjMT">Mandragora</a> (2 sp.)</li></ul></li><li>Tribe Nicandreae<ul><li><a href="/facts/Exodeconus/xKftt8Rg">Exodeconus</a> (6 sp.)</li><li><a href="/facts/Nicandra/sA4LsruE">Nicandra</a> (3 sp.)</li></ul></li><li>Tribe Solandreae<ul><li><a href="/facts/Doselia/mt6dXmJ2">Doselia</a> (4 sp.)</li><li><a href="/facts/Dyssochroma/MuFEqnCY">Dyssochroma</a> (3 sp.)</li><li><a href="/facts/Hawkesiophyton/JPD4CtIP">Hawkesiophyton</a> (3 sp.)</li><li><a href="/facts/Juanulloa/GvXTHR8D">Juanulloa</a> (8 sp.)</li><li><a href="/facts/Markea/njKCohtd">Markea</a> (14 sp.)</li><li><a href="/facts/Merinthopodium/5PG3Pom5">Merinthopodium</a> (3 sp.)</li><li><a href="/facts/Poortmannia/rJpMNpHQ">Poortmannia</a> (1 sp.)</li><li><a href="/facts/Schultesianthus/64yv4fE8">Schultesianthus</a> (8 sp.)</li><li><a href="/facts/Solandra/KyWtjAMT">Solandra</a> (10 sp.)</li><li><a href="/facts/Trianaea/StSTIj16">Trianaea</a> (3 sp.)</li></ul></li></ul></td><td>Subfamily Solanoideae (cont.)<ul><li>Tribe Physalideae<ul><li><a href="/facts/Alkekengi/NIrmGE3a">Alkekengi</a> (1 sp.)</li><li><a href="/facts/Athenaea_(plant)/4YbDMJy8">Athenaea</a> (14 sp.)</li><li><a href="/facts/Calliphysalis/2z92pvhs">Calliphysalis</a> (1 sp.)</li><li><a href="/facts/Capsicophysalis/0yNAuUVL">Capsicophysalis</a> (1 sp.)</li><li><a href="/facts/Cataracta/08NqpHBY">Cataracta</a> (1 sp.)</li><li><a href="/facts/Chamaesaracha/JpsQIeDP">Chamaesaracha</a> (12 sp.)</li><li><a href="/facts/Cuatresia/cByjkaLG">Cuatresia</a> (19 sp.)</li><li><a href="/facts/Darcyanthus/TNQXRQv6">Darcyanthus</a> (1 sp.)</li><li><a href="/facts/Deprea/4pRgdnCu">Deprea</a> (52 sp.)</li><li><a href="/facts/Discopodium/5rhaEwNR">Discopodium</a> (2 sp.)</li><li><a href="/facts/Dunalia/Byjavcxn">Dunalia</a> (4 sp.)</li><li><a href="/facts/Eriolarynx/tCapzqgl">Eriolarynx</a> (4 sp.)</li><li><a href="/facts/Iochroma/u5oEEnC4">Iochroma</a> (30 sp.)</li><li><a href="/facts/Leucophysalis/5rGultmx">Leucophysalis</a> (2 sp.)</li><li><a href="/facts/Mellissia/XpVIBHwR">Mellissia</a> (1 sp.)</li><li><a href="/facts/Nothocestrum/v3EAd73L">Nothocestrum</a> (4 sp.)</li><li><a href="/facts/Oryctes_nevadensis/J65DInpG">Orcytes</a> (9 sp.)</li><li><a href="/facts/Pantacantha/qbsQlDG7">Pantacantha</a> (1 sp.)</li><li><a href="/facts/Physalis/Cn3ThbB1">Physalis</a> (84 sp.)</li><li><a href="/facts/Quincula/3w0IwQyD">Quincula</a> (1 sp.)</li><li><a href="/facts/Saracha/q80M0fEm">Saracha</a> (5 sp.)</li><li><a href="/facts/Schraderanthus/Scq14Yjv">Schraderanthus</a> (1 sp.)</li><li><a href="/facts/Streptosolen/ox269EIz">Streptosolen</a> (1 sp.)</li><li><a href="/facts/Trozelia/aSw8FxW1">Trozelia</a> (2 sp.)</li><li>Tuberowithania (1 sp.)</li><li><a href="/facts/Tubocapsicum/r27UcXbq">Tubocapsicum</a> (1 sp.)</li><li><a href="/facts/Tzeltalia/T8qXPIj7">Tzeltalia</a> (3 sp.)</li><li><a href="/facts/Vassobia/6C95899U">Vassobia</a> (2 sp.)</li><li><a href="/facts/Withania/VILUa8fF">Withania</a> (9 sp.)</li><li><a href="/facts/Witheringia/ezmRg3X8">Witheringia</a> (19 sp.)</li></ul></li><li>Tribe Solaneae<ul><li><a href="/facts/Jaltomata/Dsf8nUBo">Jaltomata</a> (72 sp.)</li><li><a href="/facts/Solanum/UDkyXGxg">Solanum</a> (1245 sp.)</li></ul></li></ul></td></tr></tbody></table>
<h2 id="evolution">Evolution</h2>
<h3>Origin</h3>

The early evolution of Solanaceae is poorly understood in part due to the extremely sparse fossil record. Only a few dozen species in the fossil record have been firmly identified as nightshades.<a class="footnote-ref" id="fnref:39" href="#fn:39">39</a> Estimates for the origin of the family vary considerably, with <a href="/facts/Molecular_clock/wlM3LYmA">molecular clocks</a> ranging from 30.9 to 83.3 million years ago.<a class="footnote-ref" id="fnref:40" href="#fn:40">40</a> The oldest fossil nightshades however, Physalis hunickenii and <a href="/facts/Physalis_infinemundi/YomdbLko">Physalis infinemundi</a>,<a class="footnote-ref" id="fnref:41" href="#fn:41">41</a> firmly place the minimum possible age of the family at 52 million years old.<a class="footnote-ref" id="fnref:42" href="#fn:42">42</a> The existence of these fossils indicates that the family is likely significantly older than previously expected. Their position near the base of the tribe <a href="/facts/Physaleae/Byjavcxn">Physaleae</a> suggests that Solanaceae had already begun to diversify into its modern lineages by this time.<a class="footnote-ref" id="fnref:43" href="#fn:43">43</a>
Using these new fossils, a team in 2023 estimated the age of Solanaceae to be approximately 73.3 million years old. Their work proposed that after the <a href="/facts/K-Pg_mass_extinction/HNyDuEcx">K-Pg mass extinction</a>, the family began to rapidly diversify, with all subfamilies diverging from each other by 56 million years ago. The subfamily <a href="/facts/Solanoideae/4MwH5WzK">Solanoideae</a> was the last to split off from the rest, but experienced the most diversification, and now accounts for 80% of all nightshades. During the <a href="/facts/Paleocene-Eocene_thermal_maximum/QnPVzRR7">Paleocene-Eocene thermal maximum</a>, nearly all solanoid <a href="/facts/Tribe_(biology)/Uca1DPl7">tribal</a> lineages diverged from each other in only about 4 million years.<a class="footnote-ref" id="fnref:44" href="#fn:44">44</a>

<h3>Dispersion and diversification</h3>

Solanaceae originated in <a href="/facts/South_America/PhrFEMk7">South America</a>.<a class="footnote-ref" id="fnref:45" href="#fn:45">45</a><a class="footnote-ref" id="fnref:46" href="#fn:46">46</a><a class="footnote-ref" id="fnref:47" href="#fn:47">47</a> It is unique in this regard compared to other families in the order <a href="/facts/Solanales/SLSIJ2D4">Solanales</a>, which originated primarily in <a href="/facts/Africa/XkqQUP9s">Africa</a>. Its geographic isolation as a result is likely what triggered its initial diversification into a distinct lineage.<a class="footnote-ref" id="fnref:48" href="#fn:48">48</a>
From South America, nightshades rapidly colonized the rest of the <a href="/facts/Americas/hEuUTL4K">Americas</a> as they continued to diversify.<a class="footnote-ref" id="fnref:49" href="#fn:49">49</a> The spread of nightshades into the <a href="/facts/Old_World/jt02swyQ">Old World</a> happened several times throughout its history. From South America, nightshades spread west over the <a href="/facts/Pacific_Ocean/bdxgXRVM">Pacific</a> into <a href="/facts/Oceania/vzTEvsfV">Oceania</a> and east over the <a href="/facts/Atlantic_Ocean/EI4uChIb">Atlantic</a> into <a href="/facts/Africa/XkqQUP9s">Africa</a>. Nightshades reached <a href="/facts/Eurasia/KlLI9GDM">Eurasia</a> first from <a href="/facts/North_America/djNjCQIl">North America</a>, and more recently were brought over by humans during the <a href="/facts/Columbian_exchange/xaYrED8p">Columbian exchange</a>. In total, there have been about 15-20 natural dispersal events that carried Solanaceae over Earth's oceans. These events could have occurred through various methods, such as seeds being blown through wind currents or floating over the ocean's surface, or carried by migrating animals such as <a href="/facts/Bird/RJF6zwLP">birds</a>.<a class="footnote-ref" id="fnref:50" href="#fn:50">50</a><a class="footnote-ref" id="fnref:51" href="#fn:51">51</a>

<h3>Phylogeny</h3>
The relationships within Solanaceae are still largely unknown. A large study from 2013 with over 1,000 species only found weak support for many clades near the base of the tree. It found that Duckeodendron, Schizanthus (subfamily Schizanthoideae), and Goetzeoideae were among the most basal, but didn't resolve the relationships between them. Cestroideae, Schwenckioideae, and Petunioideae were found to be closer to the well supported "X=12" clade (Nicotianoideae + Solanoideae), albeit with low support. The genus <a href="/facts/Reyesia/vQYBmxYB">Reyesia</a>, usually placed in Cestroideae, was placed as the sister to Goetzeoideae.<a class="footnote-ref" id="fnref:52" href="#fn:52">52</a>
Another large study from 2023 shed more light on the early evolution of Solanaceae. It mostly corroborated and further elaborated the 2013 study, placing Schizanthus as the most basal of all nightshades, Cestroideae and Schwenkioideae as sisters, and Petunioideae as the sister to the X=12 clade. They rejected the position of Reyesia as the sister to Goetzeoideae, and placed it back into Cestroideae with both phylogenetic and morphological evidence. Like other cestroids, and indeed most other nightshades, Reyesia has radially symmetric (actinomorphic) flowers, unlike the bilaterally symmetric (zygomorphic) flowers of goetzeoids. While it had somewhat strong support the early-diverging clades, Duckeodendron was not included and the position of Goetzoideae has proved to be extremely variable compared to other studies, and they did not rule out the possibility of their placement to be incorrect.<a class="footnote-ref" id="fnref:53" href="#fn:53">53</a>

<table><tbody><tr><td>Solanaceae</td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><a href="/facts/Duckeodendron/l9nItUbL">Duckeodendron</a></td></tr><tr><td></td></tr><tr><td></td><td rowspan="2"><a href="/facts/Schizanthus/n3RYlUYF">Schizanthoideae</a> </td></tr><tr><td></td></tr><tr><td></td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2">Goetzeoideae </td></tr><tr><td></td></tr><tr><td></td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><a href="/facts/Cestroideae/086ugKao">Cestroideae</a> incl. <a href="/facts/Reyesia/vQYBmxYB">Reyesia</a> </td></tr><tr><td></td></tr><tr><td></td><td rowspan="2">Schwenckioideae </td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr><tr><td></td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><a href="/facts/Petunioideae/noe2E2d7">Petunioideae</a> </td></tr><tr><td></td></tr><tr><td>X=12 clade</td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><a href="/facts/Nicotianoideae/Tcx1sIC3">Nicotianoideae</a> </td></tr><tr><td></td></tr><tr><td></td><td rowspan="2"><a href="/facts/Solanoideae/4MwH5WzK">Solanoideae</a> </td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td><a href="/facts/Floral_symmetry/oJMuxKtL">Actinomorphic</a></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table>
<h2 id="distribution-and-habitat">Distribution and habitat</h2>

Even though members of the Solanaceae are found on all <a href="/facts/Continent/Pysd9Sw8">continents</a> except Antarctica, the greatest variety of species are found in <a href="/facts/Central_America/CXKgTuxC">Central America</a> and <a href="/facts/South_America/PhrFEMk7">South America</a>. Centers of diversity also occur in <a href="/facts/Australia/qLHpbppq">Australia</a> and <a href="/facts/Africa/XkqQUP9s">Africa</a>. Solanaceae occupy a great number of different <a href="/facts/Ecosystem/kVE8mSmR">ecosystems</a>, from <a href="/facts/Desert/Mknp4bXq">deserts</a> to <a href="/facts/Rainforest/IEY5TMEc">rainforests</a>, and are often found in the secondary vegetation that colonizes disturbed areas. In general, plants in this family are of tropical and temperate distribution.<a class="footnote-ref" id="fnref:54" href="#fn:54">54</a>

<h2 id="ecology">Ecology</h2>
The potato tuber moth (<a href="/facts/Phthorimaea_operculella/9c0mblal">Phthorimaea operculella</a>) is an oligophagous insect that prefers to feed on plants of the family Solanaceae, especially the potato plant (Solanum tuberosum). Female P. operculella use the leaves to lay their eggs and the hatched larvae will eat away at the mesophyll of the leaf. After feeding on the foliage, the larvae will then delve down and feed on the tubers and roots of the plant.<a class="footnote-ref" id="fnref:55" href="#fn:55">55</a>

<h2 id="alkaloids">Alkaloids</h2>
Hundreds of alkaloids are produced by various species of nightshades, including <a href="/facts/Solanine/Iwxa3UI5">solanine</a>, <a href="/facts/Chaconine/hHb6pRRJ">chaconine</a>, <a href="/facts/Atropine/cJQ0gL4o">atropine</a>, <a href="/facts/Tomatine/HGNyBKHD">tomatine</a>, and several more. These are used as a natural <a href="/facts/Pesticide/Uhf4KnwJ">pesticide</a> by the plant, disrupting cellular and <a href="/facts/Physiology/fh25y2pQ">physiological</a> processes in invading organisms . They are mainly targeted <a href="/facts/Insecticide/1xYL3MSD">against insects</a>, but can also function as <a href="/facts/Fungicide/c1twQBYo">fungicides</a>, <a href="/facts/Nematicide/xnJNqGxx">nematicides</a>, and <a href="/facts/Bactericide/X1XwY97r">bactericides</a>.<a class="footnote-ref" id="fnref:56" href="#fn:56">56</a> In high amounts, some alkaloids can also be toxic to humans, but others are sought after for medicinal, recreational, or culinary purposes.

<h3>Solanine</h3>

<a href="/facts/Solanine/Iwxa3UI5">Solanine</a> is a toxic <a href="/facts/Glycoalkaloid/Ko8cNWQu">glycoalkaloid</a> with a bitter taste, it has the formula C45H73NO15. It is formed by the alkaloid solanidine with a <a href="/facts/Carbohydrate/Geuuzxsf">carbohydrate</a> side chain. It is found in leaves, fruit, and tubers of various Solanaceae such as the potato and tomato. Its production is thought to be an adaptive defence strategy against herbivores. <a href="/facts/Substance_intoxication/LRnMxrKd">Substance intoxication</a> from solanine is characterized by gastrointestinal disorders (<a href="/facts/Diarrhoea/6TBHaRdA">diarrhoea</a>, <a href="/facts/Vomiting/R2vQkHqV">vomiting</a>, abdominal pain) and neurological disorders (<a href="/facts/Hallucination/5eUvLcnp">hallucinations</a> and <a href="/facts/Headache/mdmmC9DP">headache</a>). The <a href="/facts/Median_lethal_dose/PSTer8Wv">median lethal dose</a> is between 2 and 5 mg/kg of body weight. Symptoms manifest 8 to 12 hours after ingestion. The amount of these glycoalkaloids in potatoes, for example, varies significantly depending on environmental conditions during their cultivation, the length of storage, and the variety. The average glycoalkaloid concentration is 0.075 mg/g of potato.<a class="footnote-ref" id="fnref:57" href="#fn:57">57</a> Solanine has occasionally been responsible for poisonings in people who ate berries from species such as <a href="/facts/Solanum_nigrum/MGlf0uFc">Solanum nigrum</a> or <a href="/facts/Solanum_dulcamara/tQCWRW54">Solanum dulcamara</a>, or green potatoes.<a class="footnote-ref" id="fnref:58" href="#fn:58">58</a><a class="footnote-ref" id="fnref:59" href="#fn:59">59</a>

<h3>Tropanes</h3>

The term "<a href="/facts/Tropanes/0TDQJFEn">tropane</a>" comes from the genus <a href="/facts/Atropa/1WlXXWgL">Atropa</a>, named after the <a href="/facts/Moirai/GXrR5RV3">Greek Fate</a>, <a href="/facts/Atropos/v9hPyQ1w">Atropos</a>, who cut the thread of life. This nomenclature reflects its toxicity and lethality. They are bicyclic organic nitrogen compounds (<a href="/facts/IUPAC/2AV6myCK">IUPAC</a> nomenclature: 8-methyl-8-azabicyclo[3.2.1]octane), with the chemical formula of C8H15N. These alkaloids include, among others, atropine, <a href="/facts/Cocaine/rdQP4zwF">cocaine</a>, <a href="/facts/Scopolamine/Fy2S6Lgf">scopolamine</a>, and <a href="/facts/Hyoscyamine/g3LEmA86">hyoscyamine</a>. They are found in various species, such as mandrake (<a href="/facts/Mandragora_officinarum/Ch2FfzwR">Mandragora officinarum</a> and <a href="/facts/Mandragora_autumnalis/QjgPDNP6">M. autumnalis</a> ), black henbane or stinking nightshade (<a href="/facts/Hyoscyamus_niger/sy4Negv7">Hyoscyamus niger</a>), belladonna (<a href="/facts/Atropa_belladonna/30NGcju6">Atropa belladonna</a>), jimson weed or devil's snare (<a href="/facts/Datura_stramonium/N0RtGKrF">Datura stramonium</a>) and <a href="/facts/Brugmansia/TD8YtCmd">Brugmansia</a> , as well as many others in the family Solanaceae.<a class="footnote-ref" id="fnref:60" href="#fn:60">60</a> <a href="/facts/Pharmacologically/4Rx8s7Ie">Pharmacologically</a>, they are the most powerful known <a href="/facts/Anticholinergic/9mDjo23r">anticholinergics</a> in existence, meaning they inhibit the <a href="/facts/Neurological/TXYEMSYI">neurological</a> signals transmitted by the <a href="/facts/Endogenous/DFkSLkRI">endogenous</a> <a href="/facts/Neurotransmitter/LlpYFSU3">neurotransmitter</a>, <a href="/facts/Acetylcholine/v1d3wRUd">acetylcholine</a>. More commonly, they can halt many types of <a href="/facts/Allergic/dYAVosh8">allergic</a> reactions. Symptoms of overdose may include <a href="/facts/Xerostomia/aKMwe59z">dry mouth</a>, <a href="/facts/Mydriasis/19r5uMvC">dilated pupils</a>, <a href="/facts/Ataxia/suENWGS6">ataxia</a>, <a href="/facts/Urinary_retention/YvTCprKk">urinary retention</a>, <a href="/facts/Hallucination/5eUvLcnp">hallucinations</a>, <a href="/facts/Seizure/zuIrQyze">convulsions</a>, <a href="/facts/Coma/N9oQ7g7R">coma</a>, and death. Atropine, a commonly used <a href="/facts/Ophthalmology/024hdTWF">ophthalmological</a> agent, dilates the pupils and thus facilitates examination of the interior of the eye. In fact, juice from the berries of A. belladonna were used by Italian courtesans during the <a href="/facts/Renaissance/4D9Bc6KU">Renaissance</a> to exaggerate the size of their eyes by causing the dilation of their pupils ("bella donna" means "pretty woman" in Italian). Despite the extreme toxicity of the tropanes, they are useful drugs when administered in extremely small dosages. They can reverse <a href="/facts/Cholinergic/51ttI8RK">cholinergic</a> poisoning, which can be caused by overexposure to organophosphate <a href="/facts/Insecticide/1xYL3MSD">insecticides</a> and <a href="/facts/Chemical_warfare/zT4t2DGk">chemical warfare</a> agents such as <a href="/facts/Sarin/aYEK2f61">sarin</a> and <a href="/facts/VX_(nerve_agent)/71BKt6Gm">VX</a>. <a href="/facts/Scopolamine/Fy2S6Lgf">Scopolamine</a> (found in <a href="/facts/Hyoscyamus_muticus/q16ns9Ht">Hyoscyamus muticus</a> and <a href="/facts/Scopolia_carniolica/x5Nb5EQ2">Scopolia carniolica</a>), is used as an <a href="/facts/Antiemetic/GEqXzKJJ">antiemetic</a> against <a href="/facts/Motion_sickness/kBs72ygo">motion sickness</a> or for people suffering from <a href="/facts/Nausea/dyhkcbeD">nausea</a> as a result of receiving <a href="/facts/Chemotherapy/fWTLbaUT">chemotherapy</a>.<a class="footnote-ref" id="fnref:61" href="#fn:61">61</a><a class="footnote-ref" id="fnref:62" href="#fn:62">62</a> Scopolamine and hyoscyamine are the most widely used tropane alkaloids in pharmacology and medicine due to their effects on the <a href="/facts/Parasympathetic_nervous_system/eNzCI8Yu">parasympathetic nervous system</a>. Atropine has a <a href="/facts/Stimulant/nmBdAXM5">stimulant</a> effect on the <a href="/facts/Central_nervous_system/k8I6To97">central nervous system</a> and heart, whereas scopolamine has a <a href="/facts/Sedative/JWzvq66P">sedative</a> effect. These alkaloids cannot be substituted by any other class of compounds, so they are still in demand. This is one of the reasons for the development of an active field of research into the metabolism of the alkaloids, the enzymes involved, and the genes that produce them. Hyoscyamine 6-β-hydroxylase, for example, catalyses the hydroxylation of hyoscyamine that leads to the production of scopolamine at the end of the tropane's biosynthetic pathway. This enzyme has been isolated and the corresponding gene cloned from three species: H. niger, A. belladonna and B. candida.<a class="footnote-ref" id="fnref:63" href="#fn:63">63</a><a class="footnote-ref" id="fnref:64" href="#fn:64">64</a><a class="footnote-ref" id="fnref:65" href="#fn:65">65</a>

<h3>Nicotine</h3>

<a href="/facts/Nicotine/VQ6HcIrY">Nicotine</a> (<a href="/facts/IUPAC/2AV6myCK">IUPAC</a> nomenclature (S)-3-(1-methylpyrrolidin-2-yl) pyridine) is a <a href="/facts/Pyrrolidine/YPVKcwyT">pyrrolidine</a> alkaloid produced in large quantities in the <a href="/facts/Tobacco/bTD4RUPQ">tobacco</a> plant (Nicotiana tabacum). Edible Solanaceae such as eggplants, tomatoes, potatoes, and peppers also contain nicotine, but at concentrations 100,000 to 1,000,000 times less than tobacco.<a class="footnote-ref" id="fnref:66" href="#fn:66">66</a><a class="footnote-ref" id="fnref:67" href="#fn:67">67</a> Nicotine's function in a plant is to act as a defense against <a href="/facts/Herbivore/me4YFdQA">herbivores</a>, as it is a very effective <a href="/facts/Neurotoxin/I7nLuFE7">neurotoxin</a>, in particular against <a href="/facts/Insect/IgDNvnUL">insects</a>. In fact, nicotine has been used for many years as an <a href="/facts/Insecticide/1xYL3MSD">insecticide</a>, though its use is currently being replaced by synthetic molecules derived from its structure. At low concentrations, nicotine acts as a stimulant in mammals, which causes the dependency in smokers. Like the tropanes, it acts on cholinergic neurons, but with the opposite effect (it is an <a href="/facts/Agonist/nz8U5NNp">agonist</a> as opposed to an <a href="/facts/Receptor_antagonist/PuXHVNdV">antagonist</a>). It has a higher specificity for <a href="/facts/Nicotinic_acetylcholine_receptor/k2s2pPHr">nicotinic acetylcholine receptors</a> than other ACh proteins.

<h3>Capsaicin</h3>

<a href="/facts/Capsaicin/aXaaFxDp">Capsaicin</a> (<a href="/facts/IUPAC/2AV6myCK">IUPAC</a> nomenclature 8-methyl-N-vanillyl-trans-6-nonenamide) is structurally different from nicotine and the tropanes. It is found in species of the genus <a href="/facts/Capsicum/oqoEFu4F">Capsicum</a>, which includes <a href="/facts/Capsicum_frutescens/pDG6Pxtx">chilis</a> and <a href="/facts/Capsicum_chinense/16BD8xg0">habaneros</a> and it is the active ingredient that determines the <a href="/facts/Scoville_scale/Do4qGYeX">Scoville rating</a> of these spices. The compound is not noticeably toxic to humans. However, it stimulates specific pain receptors in the majority of mammals, specifically those related to the perception of heat in the <a href="/facts/Oral_mucosa/uJyDj6c0">oral mucosa</a> and other <a href="/facts/Epithelium/ecKHcHC3">epithelial tissues</a>. When capsaicin comes into contact with these mucosae, it causes a burning sensation little different from a burn caused by fire. Capsaicin affects only mammals, not birds. Pepper seeds can survive the digestive tracts of birds; their fruit becomes brightly coloured once its seeds are mature enough to germinate, thereby attracting the attention of birds that then distribute the seeds. Capsaicin extract is used to make <a href="/facts/Pepper_spray/opIwYmws">pepper spray</a>, a useful deterrent against aggressive mammals.

<h2 id="economic-importance">Economic importance</h2>

The family Solanaceae contains such important food species as the potato (<a href="/facts/Solanum_tuberosum/F8jRVExs">Solanum tuberosum</a>), the tomato (<a href="/facts/Solanum_lycopersicum/3d5Xsqw7">Solanum lycopersicum</a>), the pepper (<a href="/facts/Capsicum/oqoEFu4F">Capsicum sp</a>) and the aubergine or eggplant (<a href="/facts/Solanum_melongena/UTRtUSsd">Solanum melongena</a>). <a href="/facts/Nicotiana_tabacum/WQKhTHHK">Nicotiana tabacum</a>, originally from South America, is now cultivated throughout the world to produce tobacco.
Many solanaceas are important weeds in various parts of the world. Their importance lies in the fact that they can host pathogens or diseases of the cultivated plants, therefore their presence increases the loss of yield or the quality of the harvested product. An example of this can be seen with <a href="/facts/Acnistus_arborescens/JQPYha47">Acnistus arborescens</a> and Browalia americana that host <a href="/facts/Thrips/FEW4Njc0">thrips</a>, which cause damage to associated cultivated plants,<a class="footnote-ref" id="fnref:68" href="#fn:68">68</a> and certain species of Datura that play host to various types of virus that are later transmitted to cultivated solanaceas.<a class="footnote-ref" id="fnref:69" href="#fn:69">69</a> Some species of weeds such as, Solanum mauritianum in South Africa represent such serious ecological and economic problems that studies are being carried out with the objective of developing a biological control through the use of insects.<a class="footnote-ref" id="fnref:70" href="#fn:70">70</a>
A wide variety of plant species and their cultivars belonging to the Solanaceae are grown as ornamental trees, shrubs, annuals and herbaceous perennials<a class="footnote-ref" id="fnref:71" href="#fn:71">71</a> Examples include <a href="/facts/Brugmansia_%25C3%2597_candida/0GqakDcd">Brugmansia × candida</a> ("angel's trumpet") grown for its large pendulous trumpet-shaped flowers, or <a href="/facts/Brunfelsia_latifolia/5Aj9CgGE">Brunfelsia latifolia</a>, whose flowers are very fragrant and change colour from violet to white over a period of 3 days. Other shrub species that are grown for their attractive flowers are <a href="/facts/Lycianthes_rantonnetii/5og5AaKw">Lycianthes rantonnetii</a> (Blue Potato Bush or Paraguay Nightshade) with violet-blue flowers and <a href="/facts/Nicotiana_glauca/ooIfkwPL">Nicotiana glauca</a> ("Tree Tobacco") Other solanaceous species and genera that are grown as ornamentals are the <a href="/facts/Petunia/ASe88e60">petunia</a> (Petunia × hybrida), <a href="/facts/Lycium/4VVD4sFu">Lycium</a>, <a href="/facts/Solanum/UDkyXGxg">Solanum</a>, <a href="/facts/Cestrum/YbBzKxS5">Cestrum</a>, Calibrachoa × hybrida and <a href="/facts/Solandra/KyWtjAMT">Solandra</a>. There is even a hybrid between Petunia and Calibrachoa (which constitutes a new <a href="/facts/Nothotaxon/RJUvgany">nothogenus</a> called × Petchoa G. Boker & J. Shaw) that is being sold as an ornamental.<a class="footnote-ref" id="fnref:72" href="#fn:72">72</a> Many other species, in particular those that produce alkaloids, are used in pharmacology and medicine (<a href="/facts/Nicotiana/Bfh1JnE0">Nicotiana</a>, <a href="/facts/Hyoscyamus/JfYN6kbo">Hyoscyamus</a>, and <a href="/facts/Datura/ma7yqk7U">Datura</a>).<a class="footnote-ref" id="fnref:73" href="#fn:73">73</a>

<h3>Genomics</h3>
Many of the species belonging to this family, among them tobacco and the tomato, are <a href="/facts/Model_organisms/vNv6q4yb">model organisms</a> that are used for research into fundamental biological questions. One of the aspects of the solanaceas' <a href="/facts/Genomics/pVU3BGpL">genomics</a> is an international project that is trying to understand how the same collection of genes and proteins can give rise to a group of organisms that are so morphologically and ecologically different. The first objective of this project was to sequence the <a href="/facts/Genome/tjhWET4V">genome</a> of the tomato. In order to achieve this each of the 12 <a href="/facts/Chromosome/FRGf11FC">chromosomes</a> of the tomato's haploid genome was assigned to different sequencing centres in different countries. So chromosomes 1 and 10 were sequenced in the United States, 3 and 11 in China, 2 in <a href="/facts/Korea/cj2kDBzc">Korea</a>, 4 in Britain, 5 in India, 7 in France, 8 in Japan, 9 in Spain and 12 in Italy. The sequencing of the <a href="/facts/Mitochondria/ErHqrdJ1">mitochondrial</a> genome was carried out in Argentina and the <a href="/facts/Chloroplast/fo8U3yTx">chloroplast</a> genome was sequenced in the <a href="/facts/European_Union/6GYcR8qM">European Union</a>.<a class="footnote-ref" id="fnref:74" href="#fn:74">74</a><a class="footnote-ref" id="fnref:75" href="#fn:75">75</a>

<h2 id="notes">Notes</h2>

<h2 id="see-also">See also</h2>
<ul><li><a href="/facts/List_of_plants_poisonous_to_equines/WXM1kwgh">List of plants poisonous to equines</a></li></ul>

<ul><li>D'Arcy, William G. (1986). Solanaceae. Columbia University Press. <a href="/facts/ISBN_(identifier)/15AdSPa9">ISBN</a> 978-0-231-05780-6.</li>
<li>Stevens, P. F. (2001–2007). <a href="http://www.mobot.org/MOBOT/Research/APweb/orders/solanalesweb.htm#Solanaceae">"Solanaceae"</a>. <a href="http://www.mobot.org/MOBOT/Research/APweb/welcome.html">Angiosperm Phylogeny Website</a>, version 8, June 2007. Retrieved 2007-11-04.</li>
<li>Watson, L.; Dallwitz, M. J. <a href="https://web.archive.org/web/20071026055528/http://delta-intkey.com/angio/www/solanace.htm">"Solanaceae"</a>. <a href="http://delta-intkey.com/angio/www/index.htm">The families of flowering plants: descriptions, illustrations, identification, and information retrieval. Version: 1 June 2007.</a> Archived from <a href="http://delta-intkey.com/angio/www/solanace.htm">the original</a> on 2007-10-26. Retrieved 2007-11-04.</li>
<li>Dimitri, M. 1987. Enciclopedia Argentina de Agricultura y Jardinería. Tomo I. Descripción de plantas cultivadas. Editorial ACME S.A.C.I., Buenos Aires.</li>
<li><a href="http://www.nhm.ac.uk/research-curation/projects/solanaceaesource//">"Solanaceae Source"</a>. Retrieved 2007-11-17.</li>
<li>Hunziker, Armando T. 2001. The Genera of Solanaceae. A.R.G. Gantner Verlag K.G., Ruggell, Liechtenstein. <a href="/facts/ISBN_(identifier)/15AdSPa9">ISBN</a> 3-904144-77-4.</li>
<li><a href="https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:38525-1">https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:38525-1</a> Plants of the World Online. Retrieved 2024-01-26</li></ul>
<h2 id="further-reading">Further reading</h2>
<ul><li>Hawkes, J. G.; Lester, R. N.; Skelding, A. D. (1979). The biology and taxonomy of the Solanaceae. Academic Press, London. <a href="/facts/ISBN_(identifier)/15AdSPa9">ISBN</a> 978-0-12-333150-2.</li>
<li>D'Arcy, William G. (1986). Solanaceae. Columbia University Press. <a href="/facts/ISBN_(identifier)/15AdSPa9">ISBN</a> 978-0-231-05780-6.</li>
<li>Radford, Albert E. (1986). <a href="https://archive.org/details/fundamentalsofpl0000radf_n5o1">Fundamentals of Plant Systematics</a>. Harper & Row, Publishers, Inc. <a href="/facts/ISBN_(identifier)/15AdSPa9">ISBN</a> 978-0-06-045305-3.</li>
<li>Kubo, Ken-ichi; Paape, Timothy; Hatakeyama, Masaomi; Entani, Tetsuyuki; Takara, Akie; Kajihara, Kie; Tsukahara, Mai; Shimizu-Inatsugi, Rie; Shimizu, Kentaro K.; Takayama, Seiji (2015). <a href="https://www.zora.uzh.ch/id/eprint/104475/7/Petunia2015.pdf">"Gene duplication and genetic exchange drive the evolution of S-RNase-based self-incompatibility in Petunia"</a> (PDF). Nature Plants. 1 (1): 14005. <a href="/facts/Bibcode_(identifier)/9HtdQSGB">Bibcode</a>:<a href="https://ui.adsabs.harvard.edu/abs/2015NatPl...114005K">2015NatPl...114005K</a>. <a href="/facts/Doi_(identifier)/muM9Etpq">doi</a>:<a href="https://doi.org/10.1038%2Fnplants.2014.5">10.1038/nplants.2014.5</a>. <a href="/facts/PMID_(identifier)/JlHAvMHt">PMID</a> <a href="https://pubmed.ncbi.nlm.nih.gov/27246052">27246052</a>.
<ul><li><a href="http://www.eurekalert.org/pub_releases/2015-01/uoz-ssp010815.php">"Sophisticated system prevents self-fertilization in petunias"</a>. EurekAlert! (Press release). January 8, 2015.</li></ul></li></ul>
<h2 id="external-links">External links</h2>

<a href="/facts/Wikispecies/b2GhA7Le">Wikispecies</a> has information related to Solanaceae.

Wikimedia Commons has media related to Solanaceae.

<a href="/facts/Wikisource/TrMdylNf">Wikisource</a> has the text of the <a href="/facts/Encyclop%25C3%25A6dia_Britannica_Eleventh_Edition/mPElbCRY">1911 Encyclopædia Britannica</a> article "Nightshade".

<ul><li><a href="http://solgenomics.net">Sol Genomics Network</a></li>
<li><a href="https://web.archive.org/web/20050307011715/http://solanaceae.net/index.php">Solanaceae Network - pictures of plants</a></li>
<li><a href="http://www.solanaceaesource.org/">Solanaceae Source</a> - A worldwide taxonomic monograph of all species in the genus Solanum.</li>
<li><a href="http://www.chileflora.com/Florachilena/FloraEnglish/PIC_FAMILIES_SIMPLE_138.php">Solanaceae of Chile, by Chileflora</a>[permanent dead link]</li>
<li><a href="http://delta-intkey.com/angio/www/solanace.htm">Solanaceae</a> <a href="https://web.archive.org/web/20071026055528/http://delta-intkey.com/angio/www/solanace.htm">Archived</a> 2007-10-26 at the <a href="/facts/Wayback_Machine/nmQ3a6JC">Wayback Machine</a> in <a href="http://delta-intkey.com/angio/">L. Watson and M.J. Dallwitz (1992 onwards). The families of flowering plants: descriptions, illustrations, identification, information retrieval.</a></li>
<li><a href="https://plants.usda.gov/java/ClassificationServlet?source=display&classid=Solanaceae">Solanaceae</a> in USDA Plants Database.</li>
<li><a href="http://flowersinisrael.com/FamSolanaceae.html">Family Solanaceae</a> <a href="https://web.archive.org/web/20130618052644/http://flowersinisrael.com/FamSolanaceae.html">Archived</a> 2013-06-18 at the <a href="/facts/Wayback_Machine/nmQ3a6JC">Wayback Machine</a> Flowers in Israel</li>
<li><a href="http://www.sgn.cornell.edu">SOL Genomics Network, Universidad de Cornell</a></li>
<li><a href="https://web.archive.org/web/20050307011715/http://solanaceae.net/index.php">Imagines de various species of Solanaceae</a></li>
<li><a href="http://www.chileflora.com/Florachilena/FloraSpanish/PIC_FAMILIES_SIMPLE_138.php">Solanaceae de Chile, by Chileflora</a></li>
<li><a href="http://www.ejournal.unam.mx/cns/no69/CNS06908.pdf">Chilli: La especia del Nuevo Mundo</a> <a href="https://web.archive.org/web/20121114184110/http://www.ejournal.unam.mx/cns/no69/CNS06908.pdf">Archived</a> 2012-11-14 at the <a href="/facts/Wayback_Machine/nmQ3a6JC">Wayback Machine</a> (Article in Spanish by Germán Octavio López Riquelme regarding the biology, nutrition, culture and medical aspects of Chile.</li>
<li><a href="http://solgenomics.net/community/links/related_sites.pl">Solanaceae Resources on the Web</a></li>
<li>Jäpelt RB, Jakobsen J (2013) Vitamin D in plants: a review of occurrence, analysis, and biosynthesis. Front Plant Sci 4, No. 136 <a href="http://www.frontiersin.org/Journal/Abstract.aspx?s=907&name=plant_physiology&ART_DOI=10.3389/fpls.2013.00136">-- Note the reference to higher cholesterol levels (and consequent Vitamin D3 levels) in family Solanaceae</a></li></ul>

<h2 id="references">References</h2>

<ol>
<li id="fn:1">Lee, M. R. (June 2006). "The Solanaceae: foods and poisons". Journal of the Royal College of Physicians of Edinburgh. 36 (2): 162–169. doi:10.1177/1478271520063602021. ISSN 1478-2715. PMID 17153152. Archived from the original on 2023-03-09. Retrieved 2023-03-09. <a href="https://pubmed.ncbi.nlm.nih.gov/17153152/" target="_blank">https://pubmed.ncbi.nlm.nih.gov/17153152/</a> <a href="#fnref:1" class="footnote-back-ref">↩</a></li>
<li id="fn:2">Merriam-Webster, Merriam-Webster's Unabridged Dictionary, Merriam-Webster, archived from the original on 2020-05-25, retrieved 2016-07-25. <a href="/wiki/Merriam-Webster" target="_blank">/wiki/Merriam-Webster</a> <a href="#fnref:2" class="footnote-back-ref">↩</a></li>
<li id="fn:3">Merriam-Webster, Merriam-Webster's Unabridged Dictionary, Merriam-Webster, archived from the original on 2020-05-25, retrieved 2016-07-25. <a href="/wiki/Merriam-Webster" target="_blank">/wiki/Merriam-Webster</a> <a href="#fnref:3" class="footnote-back-ref">↩</a></li>
<li id="fn:4">Harper, Douglas. "nightshade". Online Etymology Dictionary. <a href="https://www.etymonline.com/?term=nightshade" target="_blank">https://www.etymonline.com/?term=nightshade</a> <a href="#fnref:4" class="footnote-back-ref">↩</a></li>
<li id="fn:5">da Silveira de Souza, Leonardo; Ott Andrade, Bianca; Renato Stehmann, João (7 Feb 2023). "An overview on studies of species complexes in Solanaceae". Acta Botanica Brasilica. 37. Bibcode:2023AcBBr..37E0032S. doi:10.1590/1677-941X-ABB-2023-0032. <a href="https://doi.org/10.1590%2F1677-941X-ABB-2023-0032" target="_blank">https://doi.org/10.1590%2F1677-941X-ABB-2023-0032</a> <a href="#fnref:5" class="footnote-back-ref">↩</a></li>
<li id="fn:6">Yasin J. Nasir. "Solanaceae". Flora of Pakistan. Archived from the original on 2009-05-14. Retrieved 2008-08-15. <a href="http://www.efloras.org/florataxon.aspx?flora_id=5&taxon_id=10828" target="_blank">http://www.efloras.org/florataxon.aspx?flora_id=5&taxon_id=10828</a> <a href="#fnref:6" class="footnote-back-ref">↩</a></li>
<li id="fn:7">Fujii, Kenjiro (1934). Cytologia. Botanical Institute. p. 281. <a href="https://books.google.com/books?id=If8ZAAAAIAAJ" target="_blank">https://books.google.com/books?id=If8ZAAAAIAAJ</a> <a href="#fnref:7" class="footnote-back-ref">↩</a></li>
<li id="fn:8">Hunziker, A.T. 1979: South American Solanaceae: a synoptic review. In: D'ARCY, W.G., 1979: The Biology and Taxonomy of the Solanaceae. Linn. Soc. Symp. Ser. 7: p 48-85. Linnean Soc. & Academic Press; London. <a href="/wiki/Armando_Theodoro_Hunziker" target="_blank">/wiki/Armando_Theodoro_Hunziker</a> <a href="#fnref:8" class="footnote-back-ref">↩</a></li>
<li id="fn:9">Balken, J.A. THE PLANT FAMILY SOLANACEAE:
FRUITS IN SOLANACEAE "Archived copy". Archived from the original on 2016-03-09. Retrieved 2013-08-09.{{cite web}}: CS1 maint: archived copy as title (link) <a href="https://web.archive.org/web/20160309121435/http://hvanbalken.com/fruit.html" target="_blank">https://web.archive.org/web/20160309121435/http://hvanbalken.com/fruit.html</a> <a href="#fnref:9" class="footnote-back-ref">↩</a></li>
<li id="fn:10">Armando T. Hunziker: The Genera of Solanaceae. A.R.G. Gantner Verlag K.G., Ruggell, Liechtenstein 2001. ISBN 3-904144-77-4 <a href="/wiki/ISBN_(identifier)" target="_blank">/wiki/ISBN_(identifier)</a> <a href="#fnref:10" class="footnote-back-ref">↩</a></li>
<li id="fn:11">"Family Solanaceae". WFO Plant List. December 2024. <a href="https://wfoplantlist.org/taxon/wfo-7000000654-2024-12?page=1" target="_blank">https://wfoplantlist.org/taxon/wfo-7000000654-2024-12?page=1</a> <a href="#fnref:11" class="footnote-back-ref">↩</a></li>
<li id="fn:12">Adanson, Michel (1763). Familles des plantes [Families of Plants] (in French). Missouri Botanical Gardens (published 1894). doi:10.5962/bhl.title.271. OCLC 6445968 – via Biodiversity Heritage Library. <a href="/wiki/Missouri_Botanical_Gardens" target="_blank">/wiki/Missouri_Botanical_Gardens</a> <a href="#fnref:12" class="footnote-back-ref">↩</a></li>
<li id="fn:13">Adanson, Michel (1763). Familles des plantes [Families of Plants] (in French). Missouri Botanical Gardens (published 1894). doi:10.5962/bhl.title.271. OCLC 6445968 – via Biodiversity Heritage Library. <a href="/wiki/Missouri_Botanical_Gardens" target="_blank">/wiki/Missouri_Botanical_Gardens</a> <a href="#fnref:13" class="footnote-back-ref">↩</a></li>
<li id="fn:14">"Solanaceae". Plants of the World Online. Royal Botanic Gardens, Kew. <a href="https://powo.science.kew.org/taxon/0000631-2" target="_blank">https://powo.science.kew.org/taxon/0000631-2</a> <a href="#fnref:14" class="footnote-back-ref">↩</a></li>
<li id="fn:15">"Solanaceae". Plants of the World Online. Royal Botanic Gardens, Kew. <a href="https://powo.science.kew.org/taxon/0000631-2" target="_blank">https://powo.science.kew.org/taxon/0000631-2</a> <a href="#fnref:15" class="footnote-back-ref">↩</a></li>
<li id="fn:16">"Verbascum blattaria". Plants of the World Online. Royal Botanic Gardens, Kew. <a href="https://powo.science.kew.org/taxon/810828-1" target="_blank">https://powo.science.kew.org/taxon/810828-1</a> <a href="#fnref:16" class="footnote-back-ref">↩</a></li>
<li id="fn:17">"Avicennia". Plants of the World Online. Royal Botanic Gardens, Kew. <a href="https://powo.science.kew.org/taxon/41194-1" target="_blank">https://powo.science.kew.org/taxon/41194-1</a> <a href="#fnref:17" class="footnote-back-ref">↩</a></li>
<li id="fn:18">"Crescentia cujete". Plants of the World Online. Royal Botanic Gardens, Kew. <a href="https://powo.science.kew.org/taxon/319161-2" target="_blank">https://powo.science.kew.org/taxon/319161-2</a> <a href="#fnref:18" class="footnote-back-ref">↩</a></li>
<li id="fn:19">de Jussieu, Antoine Laurent (1789). Genera Plantarum [Plant Genera] (PDF) (in Latin). Missouri Botanical Garden – via Biodiversity Heritage Library. <a href="https://ia601804.us.archive.org/15/items/mobot31753000471463/mobot31753000471463.pdf" target="_blank">https://ia601804.us.archive.org/15/items/mobot31753000471463/mobot31753000471463.pdf</a> <a href="#fnref:19" class="footnote-back-ref">↩</a></li>
<li id="fn:20">de Jussieu, Antoine Laurent (1789). Genera Plantarum [Plant Genera] (PDF) (in Latin). Missouri Botanical Garden – via Biodiversity Heritage Library. <a href="https://ia601804.us.archive.org/15/items/mobot31753000471463/mobot31753000471463.pdf" target="_blank">https://ia601804.us.archive.org/15/items/mobot31753000471463/mobot31753000471463.pdf</a> <a href="#fnref:20" class="footnote-back-ref">↩</a></li>
<li id="fn:21">Gray, Samuel Frederick (1821). A natural arrangement of British plants, etc. Vol. II. London.{{cite book}}: CS1 maint: date and year (link) <a href="https://books.google.com/books?id=-OEYAAAAYAAJ&printsec=frontcover#v=onepage&q&f=false" target="_blank">https://books.google.com/books?id=-OEYAAAAYAAJ&printsec=frontcover#v=onepage&q&f=false</a> <a href="#fnref:21" class="footnote-back-ref">↩</a></li>
<li id="fn:22">Burnett, Gilbert Thomas (1835). Outlines of Botany, etc. Vol. 2. London. <a href="https://books.google.com/books?id=K0vKL7Sh_dsC" target="_blank">https://books.google.com/books?id=K0vKL7Sh_dsC</a> <a href="#fnref:22" class="footnote-back-ref">↩</a></li>
<li id="fn:23">Maiden, J. H. (1905). "The International Rules of Botanical Nomenclature". Journal and proceedings of the Royal Society of New South Wales. 40: 74–94 – via Biodiversity Heritage Libary. <a href="https://www.biodiversitylibrary.org/part/214606" target="_blank">https://www.biodiversitylibrary.org/part/214606</a> <a href="#fnref:23" class="footnote-back-ref">↩</a></li>
<li id="fn:24">Burnett, Gilbert Thomas (1835). Outlines of Botany, etc. Vol. 2. London. <a href="https://books.google.com/books?id=K0vKL7Sh_dsC" target="_blank">https://books.google.com/books?id=K0vKL7Sh_dsC</a> <a href="#fnref:24" class="footnote-back-ref">↩</a></li>
<li id="fn:25">Burnett, Gilbert Thomas (1835). Outlines of Botany, etc. Vol. 2. London. <a href="https://books.google.com/books?id=K0vKL7Sh_dsC" target="_blank">https://books.google.com/books?id=K0vKL7Sh_dsC</a> <a href="#fnref:25" class="footnote-back-ref">↩</a></li>
<li id="fn:26">Cronk, Quentin C.B.; Bateman, Richard M.; Hawkins, Julie A. (29 January 2004). "14". Developmental Genetics and Plant Evolution. <a href="https://books.google.com/books?id=aZE3EQAAQBAJ&dq=goetzeaceae&lr=&source=gbs_navlinks_s" target="_blank">https://books.google.com/books?id=aZE3EQAAQBAJ&dq=goetzeaceae&lr=&source=gbs_navlinks_s</a> <a href="#fnref:26" class="footnote-back-ref">↩</a></li>
<li id="fn:27">Olmstead, Richard G.; Sweere, Jennifer A.; Spangler, Russell E.; Bohs, Lynn; Palmer, Jeffery D. (1999). "Phylogeny and provisional classification of the Solanaceae based on chloroplast DNA" (PDF). Solanaceae IV: 111–137. <a href="https://www.researchgate.net/profile/Tharindu-Ranasinghe-2/post/Is-there-a-complete-phylogenetic-description-of-the-Solanaceae-family/attachment/59d63db579197b807799a764/AS%3A421051545735172%401477397919618/download/PHYLOGENY+AND+PROVISIONAL+CLASSIFICATION+OF+THE+SOLANACEAE+BASED+ON+CHLOROPLAST+DNA.pdf" target="_blank">https://www.researchgate.net/profile/Tharindu-Ranasinghe-2/post/Is-there-a-complete-phylogenetic-description-of-the-Solanaceae-family/attachment/59d63db579197b807799a764/AS%3A421051545735172%401477397919618/download/PHYLOGENY+AND+PROVISIONAL+CLASSIFICATION+OF+THE+SOLANACEAE+BASED+ON+CHLOROPLAST+DNA.pdf</a> <a href="#fnref:27" class="footnote-back-ref">↩</a></li>
<li id="fn:28">Ganaie, Masood Majaz; Raja, Vaseem; Reshi, Zaffar Ahmad; Verma, Vijeshwer (12 July 2018). "Family Solanaceae: Taxonomy and modern trends" (PDF). Annals of Plant Sciences: 2403–2414. ISSN 2287-688X. <a href="https://d1wqtxts1xzle7.cloudfront.net/67313064/pdf-libre.pdf?1620912311=&response-content-disposition=inline%3B+filename%3DFamily_Solanaceae_Taxonomy_and_modern_tr.pdf&Expires=1748529402&Signature=AqBA3SKyH7vWeDHBogQWoQy2Nb9r9APQbL3ql3oC7zz9zNXhmtjOvhxXpYtLRDhR3lyDw8Tz40GWNCDU4~Mz1F-PIkn25XZ4MgpQfrhsJba8YSsmEIADGGgKEOZCAlv7H2n0EVCCwaHSTSDrfmtzeX6SahDlDUpyYFy25OpYsvcmGCWMA3HRd4cyPbPEsQvdyuTlnRBVC68H1X9cjRjSiZpubp~-27EkQXUs-u1RV1MUc7SHoxEGb2-FHMNffjoU~d8iMGkuvoH5o8xpHA93Rsvru8Zyt0K5Gva2QLBQfNek4Svo2bd~DkBbv5El0EsIde0-cCLjUpEEVr1Ig08Uxw__&Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA" target="_blank">https://d1wqtxts1xzle7.cloudfront.net/67313064/pdf-libre.pdf?1620912311=&response-content-disposition=inline%3B+filename%3DFamily_Solanaceae_Taxonomy_and_modern_tr.pdf&Expires=1748529402&Signature=AqBA3SKyH7vWeDHBogQWoQy2Nb9r9APQbL3ql3oC7zz9zNXhmtjOvhxXpYtLRDhR3lyDw8Tz40GWNCDU4~Mz1F-PIkn25XZ4MgpQfrhsJba8YSsmEIADGGgKEOZCAlv7H2n0EVCCwaHSTSDrfmtzeX6SahDlDUpyYFy25OpYsvcmGCWMA3HRd4cyPbPEsQvdyuTlnRBVC68H1X9cjRjSiZpubp~-27EkQXUs-u1RV1MUc7SHoxEGb2-FHMNffjoU~d8iMGkuvoH5o8xpHA93Rsvru8Zyt0K5Gva2QLBQfNek4Svo2bd~DkBbv5El0EsIde0-cCLjUpEEVr1Ig08Uxw__&Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA</a> <a href="#fnref:28" class="footnote-back-ref">↩</a></li>
<li id="fn:29">Johnston, Ivan M. (1936). "A Study of the Nolanaceae". Proceedings of the American Academy of Arts and Sciences. 71 (1): 1–87. doi:10.2307/20023213. ISSN 0199-9818. <a href="https://www.jstor.org/stable/20023213" target="_blank">https://www.jstor.org/stable/20023213</a> <a href="#fnref:29" class="footnote-back-ref">↩</a></li>
<li id="fn:30">Carlquist, Sherwin (1988-01-01). "Wood Anatomy and Relationships of Duckeodendraceae and Goetzeaceae". IAWA Journal. 9 (1): 3–12. doi:10.1163/22941932-90000460. ISSN 0928-1541. <a href="https://brill.com/view/journals/iawa/9/1/article-p3_1.xml" target="_blank">https://brill.com/view/journals/iawa/9/1/article-p3_1.xml</a> <a href="#fnref:30" class="footnote-back-ref">↩</a></li>
<li id="fn:31">Carlquist, Sherwin (1988-01-01). "Wood Anatomy and Relationships of Duckeodendraceae and Goetzeaceae". IAWA Journal. 9 (1): 3–12. doi:10.1163/22941932-90000460. ISSN 0928-1541. <a href="https://brill.com/view/journals/iawa/9/1/article-p3_1.xml" target="_blank">https://brill.com/view/journals/iawa/9/1/article-p3_1.xml</a> <a href="#fnref:31" class="footnote-back-ref">↩</a></li>
<li id="fn:32">Zona, Scott (1989-01-01). "Leaf Anatomy of the Goetzeaceae". Aliso: A Journal of Systematic and Floristic Botany. 12 (2): 303–312. doi:10.5642/aliso.19891202.07. ISSN 0065-6275. <a href="https://scholarship.claremont.edu/aliso/vol12/iss2/7" target="_blank">https://scholarship.claremont.edu/aliso/vol12/iss2/7</a> <a href="#fnref:32" class="footnote-back-ref">↩</a></li>
<li id="fn:33">Young, Andrew D.; Gillung, Jessica P. (2020). "Phylogenomics — principles, opportunities and pitfalls of big-data phylogenetics". Systematic Entomology. 45 (2): 225–247. doi:10.1111/syen.12406. ISSN 1365-3113. <a href="https://onlinelibrary.wiley.com/doi/abs/10.1111/syen.12406" target="_blank">https://onlinelibrary.wiley.com/doi/abs/10.1111/syen.12406</a> <a href="#fnref:33" class="footnote-back-ref">↩</a></li>
<li id="fn:34">Olmstead, Richard G.; Sweere, Jennifer A.; Spangler, Russell E.; Bohs, Lynn; Palmer, Jeffery D. (1999). "Phylogeny and provisional classification of the Solanaceae based on chloroplast DNA" (PDF). Solanaceae IV: 111–137. <a href="https://www.researchgate.net/profile/Tharindu-Ranasinghe-2/post/Is-there-a-complete-phylogenetic-description-of-the-Solanaceae-family/attachment/59d63db579197b807799a764/AS%3A421051545735172%401477397919618/download/PHYLOGENY+AND+PROVISIONAL+CLASSIFICATION+OF+THE+SOLANACEAE+BASED+ON+CHLOROPLAST+DNA.pdf" target="_blank">https://www.researchgate.net/profile/Tharindu-Ranasinghe-2/post/Is-there-a-complete-phylogenetic-description-of-the-Solanaceae-family/attachment/59d63db579197b807799a764/AS%3A421051545735172%401477397919618/download/PHYLOGENY+AND+PROVISIONAL+CLASSIFICATION+OF+THE+SOLANACEAE+BASED+ON+CHLOROPLAST+DNA.pdf</a> <a href="#fnref:34" class="footnote-back-ref">↩</a></li>
<li id="fn:35">Santiago-Valentin, Eugenio; Olmstead, Richard G. (2003). "Phylogenetics of the Antillean Goetzeoideae (Solanaceae) and Their Relationships within the Solanaceae Based on Chloroplast and ITS DNA Sequence Data". Systematic Botany. 28 (2): 452–460. ISSN 0363-6445. <a href="https://www.jstor.org/stable/3094013" target="_blank">https://www.jstor.org/stable/3094013</a> <a href="#fnref:35" class="footnote-back-ref">↩</a></li>
<li id="fn:36">Särkinen, Tiina; Bohs, Lynn; Olmstead, Richard G.; Knapp, Sandra (30 September 2013). "A phylogenetic framework for evolutionary study of the nightshades (Solanaceae): a dated 1000-tip tree". BMC evolutionary biology. 13. doi:10.1186/1471-2148-13-214. ISSN 1471-2148. PMC 3850475. PMID 24283922.{{cite journal}}: CS1 maint: unflagged free DOI (link) <a href="https://pmc.ncbi.nlm.nih.gov/articles/PMC3850475/" target="_blank">https://pmc.ncbi.nlm.nih.gov/articles/PMC3850475/</a> <a href="#fnref:36" class="footnote-back-ref">↩</a></li>
<li id="fn:37">Atrichodendron and Tuberowithania are not listed under Solanaceae in the WFO Plant List, but are in their database.[33][34] While Tuberowithania is a nightshade;[35] a member of the tribe Physalideae,[36] Atrichodendron may be a member of Boraginaceae and is not included below.[37] <a href="#fnref:37" class="footnote-back-ref">↩</a></li>
<li id="fn:38">"Solanaceae". WFO Plant List. <a href="https://wfoplantlist.org/taxon/wfo-7000000654-2024-12?page=1" target="_blank">https://wfoplantlist.org/taxon/wfo-7000000654-2024-12?page=1</a> <a href="#fnref:38" class="footnote-back-ref">↩</a></li>
<li id="fn:39">Deanna, Rocío; Wilf, Peter; Gandolfo, Maria (28 November 2020). "New physaloid fruit-fossil species from early Eocene South America". American Journal of Botany. 107 (12): 1749–1762. doi:10.1002/ajb2.1565. PMID 33247843. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:39" class="footnote-back-ref">↩</a></li>
<li id="fn:40">Huang, Jie; et al. (10 July 2023). "Nuclear phylogeny and insights into whole-genome duplications and reproductive development of Solanaceae plants". Plant Communications. 4 (4). Bibcode:2023PlCom...400595H. doi:10.1016/j.xplc.2023.100595. PMC 10363554. PMID 36966360. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10363554" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10363554</a> <a href="#fnref:40" class="footnote-back-ref">↩</a></li>
<li id="fn:41">Deanna, Rocío; Wilf, Peter; Gandolfo, Maria (28 November 2020). "New physaloid fruit-fossil species from early Eocene South America". American Journal of Botany. 107 (12): 1749–1762. doi:10.1002/ajb2.1565. PMID 33247843. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:41" class="footnote-back-ref">↩</a></li>
<li id="fn:42">Särkinen, Tiina; Kottner, Sören; Stuppy, Wolfgang; Ahmed, Farah; Knapp, Sandra (12 February 2018). "A new commelinid monocot seed fossil from the early Eocene previously identified as Solanaceae". American Journal of Botany. 105 (1): 95–107. doi:10.1002/ajb2.1009. PMID 29532926. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:42" class="footnote-back-ref">↩</a></li>
<li id="fn:43">Deanna, Rocío; Wilf, Peter; Gandolfo, Maria (28 November 2020). "New physaloid fruit-fossil species from early Eocene South America". American Journal of Botany. 107 (12): 1749–1762. doi:10.1002/ajb2.1565. PMID 33247843. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:43" class="footnote-back-ref">↩</a></li>
<li id="fn:44">Huang, Jie; et al. (10 July 2023). "Nuclear phylogeny and insights into whole-genome duplications and reproductive development of Solanaceae plants". Plant Communications. 4 (4). Bibcode:2023PlCom...400595H. doi:10.1016/j.xplc.2023.100595. PMC 10363554. PMID 36966360. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10363554" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10363554</a> <a href="#fnref:44" class="footnote-back-ref">↩</a></li>
<li id="fn:45">Dupin, Julia; et al. (26 November 2016). "Bayesian estimation of the global biogeographical history of the Solanaceae". Journal of Biogeography. 44 (4): 887–899. doi:10.1111/jbi.12898. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:45" class="footnote-back-ref">↩</a></li>
<li id="fn:46">Olmstead, Richard (24 October 2012). "Phylogeny and biogeography in Solanaceae, Verbenaceae and Bignoniaceae: a comparison of continental and intercontinental diversification patterns". Botanical Journal of the Linnean Society. 171: 80–102. doi:10.1111/j.1095-8339.2012.01306.x. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:46" class="footnote-back-ref">↩</a></li>
<li id="fn:47">Deanna, Rocío; Wilf, Peter; Gandolfo, Maria (28 November 2020). "New physaloid fruit-fossil species from early Eocene South America". American Journal of Botany. 107 (12): 1749–1762. doi:10.1002/ajb2.1565. PMID 33247843. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:47" class="footnote-back-ref">↩</a></li>
<li id="fn:48">Olmstead, Richard (24 October 2012). "Phylogeny and biogeography in Solanaceae, Verbenaceae and Bignoniaceae: a comparison of continental and intercontinental diversification patterns". Botanical Journal of the Linnean Society. 171: 80–102. doi:10.1111/j.1095-8339.2012.01306.x. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:48" class="footnote-back-ref">↩</a></li>
<li id="fn:49">Dupin, Julia; et al. (26 November 2016). "Bayesian estimation of the global biogeographical history of the Solanaceae". Journal of Biogeography. 44 (4): 887–899. doi:10.1111/jbi.12898. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:49" class="footnote-back-ref">↩</a></li>
<li id="fn:50">Dupin, Julia; et al. (26 November 2016). "Bayesian estimation of the global biogeographical history of the Solanaceae". Journal of Biogeography. 44 (4): 887–899. doi:10.1111/jbi.12898. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:50" class="footnote-back-ref">↩</a></li>
<li id="fn:51">Olmstead, Richard (24 October 2012). "Phylogeny and biogeography in Solanaceae, Verbenaceae and Bignoniaceae: a comparison of continental and intercontinental diversification patterns". Botanical Journal of the Linnean Society. 171: 80–102. doi:10.1111/j.1095-8339.2012.01306.x. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:51" class="footnote-back-ref">↩</a></li>
<li id="fn:52">Särkinen, Tiina; Bohs, Lynn; Olmstead, Richard G.; Knapp, Sandra (30 September 2013). "A phylogenetic framework for evolutionary study of the nightshades (Solanaceae): a dated 1000-tip tree". BMC evolutionary biology. 13. doi:10.1186/1471-2148-13-214. ISSN 1471-2148. PMC 3850475. PMID 24283922.{{cite journal}}: CS1 maint: unflagged free DOI (link) <a href="https://pmc.ncbi.nlm.nih.gov/articles/PMC3850475/" target="_blank">https://pmc.ncbi.nlm.nih.gov/articles/PMC3850475/</a> <a href="#fnref:52" class="footnote-back-ref">↩</a></li>
<li id="fn:53">Huang, Jie; et al. (10 July 2023). "Nuclear phylogeny and insights into whole-genome duplications and reproductive development of Solanaceae plants". Plant Communications. 4 (4). Bibcode:2023PlCom...400595H. doi:10.1016/j.xplc.2023.100595. PMC 10363554. PMID 36966360. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10363554" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10363554</a> <a href="#fnref:53" class="footnote-back-ref">↩</a></li>
<li id="fn:54">Yang, Fan; Han, Songxue; Zhang, Yangxin; Chen, Xiangxiang; Gai, Wenxian; Zhao, Tao (19 October 2024). "Phylogenomic Analysis and Functional Characterization of the APETALA2/Ethylene-Responsive Factor Transcription Factor Across Solanaceae". International Journal of Molecular Sciences. 25 (20): 11247. doi:10.3390/ijms252011247. PMC 11508751. PMID 39457030. ProQuest 3120646080. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11508751" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11508751</a> <a href="#fnref:54" class="footnote-back-ref">↩</a></li>
<li id="fn:55">Varela, L. G.; Bernays, E. A. (1988-07-01). "Behavior of newly hatched potato tuber moth larvae,Phthorimaea operculella Zell. (Lepidoptera: Gelechiidae), in relation to their host plants". Journal of Insect Behavior. 1 (3): 261–275. Bibcode:1988JIBeh...1..261V. doi:10.1007/BF01054525. <a href="/wiki/Bibcode_(identifier)" target="_blank">/wiki/Bibcode_(identifier)</a> <a href="#fnref:55" class="footnote-back-ref">↩</a></li>
<li id="fn:56">Chowański, Szymon (22 February 2016). "A Review of Bioinsecticidal Activity of Solanaceae Alkaloids". Toxins. 8 (3): 60. doi:10.3390/toxins8030060. PMC 4810205. PMID 26938561. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4810205" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4810205</a> <a href="#fnref:56" class="footnote-back-ref">↩</a></li>
<li id="fn:57">Zeiger, E. 1998. Solanine and Chaconine. Review of Toxicological Literature. Integrated Laboratory Systems, USA."Archived copy" (PDF). Archived from the original (PDF) on 2011-11-15. Retrieved 2011-11-15.{{cite web}}: CS1 maint: archived copy as title (link) <a href="https://web.archive.org/web/20111115131818/http://ntp.niehs.nih.gov/ntp/htdocs/Chem_Background/ExSumPdf/ChaconineSolanine.pdf" target="_blank">https://web.archive.org/web/20111115131818/http://ntp.niehs.nih.gov/ntp/htdocs/Chem_Background/ExSumPdf/ChaconineSolanine.pdf</a> <a href="#fnref:57" class="footnote-back-ref">↩</a></li>
<li id="fn:58">"Solanine poisoning". Br Med J. 2 (6203): 1458–1459. 1979-12-08. doi:10.1136/bmj.2.6203.1458-a. PMC 1597169. PMID 526812. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1597169" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1597169</a> <a href="#fnref:58" class="footnote-back-ref">↩</a></li>
<li id="fn:59">Alexander RF, Forbes GB, Hawkins ES (1948-09-11). "A Fatal Case of Solanine Poisoning". Br Med J. 2 (4575): 518. doi:10.1136/bmj.2.4575.518. PMC 2091497. PMID 18881287. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2091497" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2091497</a> <a href="#fnref:59" class="footnote-back-ref">↩</a></li>
<li id="fn:60">Griffin WJ, Lin GD (March 2000). "Chemotaxonomy and geographical distribution of tropane alkaloids". Phytochemistry. 53 (6): 623–637. Bibcode:2000PChem..53..623G. doi:10.1016/S0031-9422(99)00475-6. PMID 10746874. <a href="/wiki/Bibcode_(identifier)" target="_blank">/wiki/Bibcode_(identifier)</a> <a href="#fnref:60" class="footnote-back-ref">↩</a></li>
<li id="fn:61">Sneden, A. The tropane alkaloids. Medicinal Chemistry and Drug Design. Virginia Commonwealth University "Archived copy" (PDF). Archived from the original (PDF) on 2007-09-27. Retrieved 2007-09-27.{{cite web}}: CS1 maint: archived copy as title (link)[unreliable source?] <a href="https://web.archive.org/web/20070927010636/http://www.people.vcu.edu/~asneden/tropane%20alkaloids.pdf" target="_blank">https://web.archive.org/web/20070927010636/http://www.people.vcu.edu/~asneden/tropane%20alkaloids.pdf</a> <a href="#fnref:61" class="footnote-back-ref">↩</a></li>
<li id="fn:62">Evans, W.C. 1979. Tropane alkaloids of the Solanaceae. En: HAWKES, LESTER and SHELDING (eds.). The biology and taxonomy of the Solanaceae. Linn. Soc. Symp. Ser. 7:241-254. Linnean Soc. & Academic Press., London. <a href="#fnref:62" class="footnote-back-ref">↩</a></li>
<li id="fn:63">Matsuda, Jun; Okabe, Souichi; Hashimoto, Takashi; Yamada, Yasuyuki (1991). "Molecular cloning of hyoscyamine 6β-hydroxylase, a 2-oxoglutarate-dependent dioxygenase, from cultured roots of Hyoscyamus niger". The Journal of Biological Chemistry. 266 (15): 9460–9464. doi:10.1016/S0021-9258(18)92843-7. PMID 2033047. <a href="https://doi.org/10.1016%2FS0021-9258%2818%2992843-7" target="_blank">https://doi.org/10.1016%2FS0021-9258%2818%2992843-7</a> <a href="#fnref:63" class="footnote-back-ref">↩</a></li>
<li id="fn:64">Rocha, Pedro; Stenzel, Olaf; Parr, Adrian; Walton, Nicholas; Christou, Paul; Dräger, Birgit; Leech, Mark J (June 2002). "Functional expression of tropinone reductase I (trI) and hyoscyamine-6β-hydroxylase (h6h) from Hyoscyamus niger in Nicotiana tabacum". Plant Science. 162 (6): 905–913. Bibcode:2002PlnSc.162..905R. doi:10.1016/S0168-9452(02)00033-X. <a href="/wiki/Bibcode_(identifier)" target="_blank">/wiki/Bibcode_(identifier)</a> <a href="#fnref:64" class="footnote-back-ref">↩</a></li>
<li id="fn:65">Cardillo, Alejandra B.; Giulietti, Ana M.; Marconi, Patricia L. (June 2006). "Analysis and sequencing of h6hmRNA, last enzyme in the tropane alkaloids pathway from anthers and hairy root cultures of Brugmansia candida (Solanaceae)". Electronic Journal of Biotechnology. 9 (3). <a href="https://www.ejbiotechnology.info/index.php/ejbiotechnology/article/view/v9n3-15" target="_blank">https://www.ejbiotechnology.info/index.php/ejbiotechnology/article/view/v9n3-15</a> <a href="#fnref:65" class="footnote-back-ref">↩</a></li>
<li id="fn:66">Siegmund, Barbara; Leitner, Erich; Pfannhauser, Werner (1999-07-23). "Determination of the Nicotine Content of Various Edible Nightshades (Solanaceae) and Their Products and Estimation of the Associated Dietary Nicotine Intake". J. Agric. Food Chem. 47 (8): 3113–3120. Bibcode:1999JAFC...47.3113S. doi:10.1021/jf990089w. PMID 10552617. <a href="/wiki/Bibcode_(identifier)" target="_blank">/wiki/Bibcode_(identifier)</a> <a href="#fnref:66" class="footnote-back-ref">↩</a></li>
<li id="fn:67">Moldoveanu, Serban C.; Scott, Wayne A.; Lawson, Darlene M. (April 2016). "Nicotine Analysis in Several Non-Tobacco Plant Materials". Beiträge zur Tabakforschung International/Contributions to Tobacco Research. 27 (2): 54–59. doi:10.1515/cttr-2016-0008. <a href="https://doi.org/10.1515%2Fcttr-2016-0008" target="_blank">https://doi.org/10.1515%2Fcttr-2016-0008</a> <a href="#fnref:67" class="footnote-back-ref">↩</a></li>
<li id="fn:68">Masís, Carlos E.; Madrigal, Rebeca (1994). "Lista preliminar de malezas hospedantes de trips (thysanoptera) que dañan al (Chrysanthemum morifolium) en el valle central de Costa Rica" [Preliminary list of thrip (Thysanoptera) weed hosts (Chrysanthemum morifolium) in the Central Valley of Costa Rica] (PDF). Agronomía costarricense (in Spanish). 18 (1): 99–101. <a href="https://www.mag.go.cr/rev_agr/v18n01_099.pdf" target="_blank">https://www.mag.go.cr/rev_agr/v18n01_099.pdf</a> <a href="#fnref:68" class="footnote-back-ref">↩</a></li>
<li id="fn:69">Ormeño, Juan; Sepúlveda, Paulina; Rojas, Ricardo; Araya, Jaime E (December 2006). "Datura Genus Weeds as an Epidemiological Factor of Alfalfa mosaic virus (AMV), Cucumber mosaic virus (CMV), and Potato virus Y (PVY) on Solanaceus Crops". Agricultura Técnica. 66 (4). doi:10.4067/S0365-28072006000400001. <a href="https://doi.org/10.4067%2FS0365-28072006000400001" target="_blank">https://doi.org/10.4067%2FS0365-28072006000400001</a> <a href="#fnref:69" class="footnote-back-ref">↩</a></li>
<li id="fn:70">Pedrosa-Macedo, José H.; Olckers, Terry; Vitorino, Marcelo D.; Caxambu, Marcelo G. (September 2003). "Phytophagous arthropods associated with Solanum mauritianum Scopoli (Solanaceae) in the first Plateau of Paraná, Brazil: a cooperative project on biological control of weeds between Brazil and South Africa". Neotropical Entomology. 32 (3). doi:10.1590/S1519-566X2003000300026. <a href="https://doi.org/10.1590%2FS1519-566X2003000300026" target="_blank">https://doi.org/10.1590%2FS1519-566X2003000300026</a> <a href="#fnref:70" class="footnote-back-ref">↩</a></li>
<li id="fn:71">"Arboles ornamentales cultivados en España. Solanáceas". Archived from the original on 2015-09-23. Retrieved 2013-08-09. <a href="http://www.arbolesornamentales.com/Solanaceae.htm" target="_blank">http://www.arbolesornamentales.com/Solanaceae.htm</a> <a href="#fnref:71" class="footnote-back-ref">↩</a></li>
<li id="fn:72">Shaw, J. M. H. (2007). "A new hybrid genus for Calibrachoa × Petunia (Solanaceae)" (PDF). Hanburyana. 2: 50–51. <a href="https://www.rhs.org.uk/about-us/pdfs/publications/hanburyana/vol-2-may-2007/a-new-hybrid-genus-for-calibrachoa-x-petunia.pdf" target="_blank">https://www.rhs.org.uk/about-us/pdfs/publications/hanburyana/vol-2-may-2007/a-new-hybrid-genus-for-calibrachoa-x-petunia.pdf</a> <a href="#fnref:72" class="footnote-back-ref">↩</a></li>
<li id="fn:73">Fatur, Karsten (June 2020). "'Hexing Herbs' in Ethnobotanical Perspective: A Historical Review of the Uses of Anticholinergic Solanaceae Plants in Europe". Economic Botany. 74 (2): 140–158. Bibcode:2020EcBot..74..140F. doi:10.1007/s12231-020-09498-w. <a href="/wiki/Bibcode_(identifier)" target="_blank">/wiki/Bibcode_(identifier)</a> <a href="#fnref:73" class="footnote-back-ref">↩</a></li>
<li id="fn:74">"International Tomato Sequencing Project Home". Archived from the original on 2009-04-27. Retrieved 2013-08-09. <a href="http://www.sgn.cornell.edu/about/tomato_sequencing.pl" target="_blank">http://www.sgn.cornell.edu/about/tomato_sequencing.pl</a> <a href="#fnref:74" class="footnote-back-ref">↩</a></li>
<li id="fn:75">"International Solanaceae Genomics Project (SOL), Systems Approach to Diversity and Adaptation". Archived from the original on 2014-08-26. Retrieved 2013-08-09. <a href="http://www.sgn.cornell.edu/solanaceae-project/index.pl" target="_blank">http://www.sgn.cornell.edu/solanaceae-project/index.pl</a> <a href="#fnref:75" class="footnote-back-ref">↩</a></li>
</ol>

Solanaceae open-in-new

Solanaceae